Range provided an operations update. As announced previously, fourth quarter production volumes averaged 541.0 Mmcfe net per day, an 18% increase over the prior-year period and 8% higher than third quarter 2010. The record production marked the Company's 32nd consecutive quarter of sequential production growth. Production was 76% natural gas, 18% natural gas liquids (NGLs) and 6% crude oil. Targeted drilling to the liquids-rich portion of the Marcellus Shale play in Pennsylvania, as well as the Midcontinent and Permian Basin regions drove the production growth. Fourth quarter 2010 production was 24% NGLs and crude oil versus 18% for fourth quarter of 2009. For the year, production increased 14%, despite property sales. Excluding the property sales, production would have grown 19%.
Fourth quarter drilling expenditures of $253 million funded the drilling of 90 (64 net) wells and 7 (7 net) recompletions. A 95% success rate was achieved. For the full-year, 237 (167 net) wells were successfully drilled and are now on production, while 111 (81 net) wells are currently in various stages of completion or waiting on pipeline connection. For the year, capital expenditures totaled just under $1.1 billion or $100 million less than budgeted.
Commenting on the announcement, John Pinkerton, Range's Chairman and CEO, said, "We ended 2010 on a very strong note, having completed the best drilling program in our history. Our 840% drill-bit reserve replacement and $0.71 all-in finding and development cost far exceeded expectations, reaching all-time records. This was accomplished even though we under spent our capital budget by approximately $100 million. 2011 greeted us with some of the toughest winter weather we have seen in many years. While Range and the industry experienced significant shut-ins due to freezing weather, our operating teams are continuing to execute by bringing on some spectacular new wells in 2011. The well results reflect the quality of our drilling inventory and the quality of our technical team. We are well-positioned to continue our streak of double digit per share production and reserve growth with one of the industry's lowest cost structures."
Range's Marcellus production for December 2010 averaged more than 200 Mmcfe per day net. Approximately 71% of the production was natural gas and 29% was NGLs and condensate. During the fourth quarter, the Marcellus division brought online 11 horizontal wells in southwestern Pennsylvania, nine of which were located in the liquids-rich wet area of the play. Based on initial production results, we expect the average estimated ultimate recovery (EUR) of these 11 wells to exceed our average reserve estimate of 5.0 Bcfe per well for the southwestern portion of Pennsylvania. At the end of the year, Range had drilled 52 horizontal wells that were waiting on completion and 15 wells waiting on pipeline connection.
Last week, Range brought on approximately 70 Mmcfe per day net of new production that, at year-end, was waiting on completion and/or pipeline connection. This included three wells in the liquid-rich area of the southwest with a collective initial production of 26.1 (22 net) Mmcf per day of natural gas and 1,373 (1,167 net) barrels of NGLs and condensate per day or 29 Mmcfe per day net. In the southwest area, Range curtailed some of its lower Btu gas to commence production from these wells in the more liquid-rich area. The curtailed production will be brought back on as we continue to build out the pipeline system in 2011. The initial phase of our Lycoming County gathering system came online February 16th. Current production from that area is 45 (39 net) Mmcf per day as we have five wells now on production, averaging 9.0 (7.9 net) Mmcf per day per well. Range plans to bring online 20 additional wells in Lycoming County by the end of the third quarter.
Earlier this month, Range completed a significant Marcellus step-out well in the southwest, which tested at 18.6 Mmcfe per day on a five-day test. Current production in the Marcellus is approximately 260 Mmcfe per day net, from both the southwest and northeast areas. Given excellent drilling results and the exceptional work of our technical team, Range is well on schedule to achieve its Marcellus Shale year-end 2011 production exit rate target of 400 Mmcfe per day net.
Providing additional resource potential from our current acreage position in this region is a variety of shale formations that lie above the Marcellus formation, known as the Upper Devonian shales. We believe that a majority of our acreage in the southwest portion of the play is prospective for the Upper Devonian shales. Earlier this month, Range announced that it believes that the Upper Devonian Shale formation on its existing acreage contains an unrisked resource potential of 10 to 14 Tcfe. Range also announced the seven-day initial production rate for the first Utica Shale horizontal completion in the basin to be 4.4 Mmcf per day. The Utica Shale formation lies below the Marcellus Shale formation.
During the fourth quarter 2010, Range's Appalachian division drilled a total of 51 (26 net) wells, continuing its successful development of tight gas sand, coal bed methane and horizontal drilling projects in Virginia. The division averaged four rigs running during the quarter and drilled 11 (6 net) vertical tight gas sand wells, 26 (12 net) coal bed methane wells and 8 (3 net) horizontal wells in Nora. In addition, on the recently acquired Nora extension property, the division drilled 5 (5 net) tight gas sand wells and 1 (1 net) horizontal Huron Shale well. The horizontal Huron Shale well was a significant step-out well drilled almost 25 miles from the prior Nora field horizontal wells. Since this area has only been sparsely drilled, reserves here are virgin pressured with no depletion from prior drilling. The step-out well tested at 2.6 Mmcf per day which is materially higher than the typical Nora horizontal well. Also in the quarter, Range performed five recompletions of behind-pipe pays on wells in the Nora extension area with positive results. Based on this success, 30 additional wells are planned for recompletion in 2011.
The Midcontinent division continued development of liquids-rich plays in the fourth quarter of 2010. Two additional horizontal Mississippian Lime wells were completed for a combined average daily rate of 807 barrels of oil, 298 barrels of NGL's and 1.3 Mmcf or 1,314 (1,035 net) Boe per day at depths of 5,000 feet. In the Ardmore Basin horizontal Woodford play we added three wells, which are currently in the completion phase and are expected to commence sales next month. One operated rig will remain active throughout the year, while non-operated activity is expected to add 1 to 2 additional rigs to the area. Non-operated activity will also drive Range's participation in the Woodford 'Cana' Shale play of the Anadarko Basin where we control approximately 80,000 (42,000) net legacy acres that are all held by production. For its first Cana Shale well, Range has joined for a 10% working interest, in a Devon Energy operated well in Blaine County, Oklahoma. The well has been drilled to total depth and completion operations have commenced. Horizontal drilling activity in the Texas Panhandle has also achieved another milestone. In the fourth quarter, Range completed the industry's first successful horizontal St. Louis Lime well which tested at rates of 360 barrels of oil, 543 barrels of NGL's and 13.8 Mmcf or 19.2 (5.8 net) Mmcfe per day net.
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